Two species of the genus Conomodulus Landau, Vermeij & Reich, 2014 are described as new from the Middle Miocene Langhian beds from Wonosari (Indonesia, Java, Yogyakarta). Recent advances in modulid taxonomy based solely on shell characters are highlighted, both at generic level and at species level, with almost 40% of known modulid species described since the year 2000. We await molecular data to test the present generic/species concepts within the family.

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Acta Malacologica Inquisitionis
Staff publications

Landau, B., & A. E. Breitenberger. (2025). Middle Miocene Modulidae (Gastropoda: Cerithioidea) from Indonesia, Java, Yogyakarta, near Wonosari. Acta Malacologica Inquisitionis, 2(1).