Studies on Lejeuneaceae subfam. Ptychanthoideae VI. A revision of Schiffneriolejeunea sect. Saccatae from Asia

The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.