Caudalejeunea grolleana Gradst. spec. nov. from Madagascar is referred to this genus with some doubt because of the absence of gemmiparous branches. Ptychocoleus cristilobus (Steph.) Steph. from S. E. Asia has gemmiparous branches and therefore is a true Caudalejeunea: C. cristiloba (Steph.) comb. nov. This species is remarkable by its complicated ciliate leaflobule and by its polystratose rhizoid-disc. Two subspecies are distinguished: ssp. cristiloba from Burma, Andaman Is., Thailand, and Singapore, and ssp. samoana (Steph.) comb. nov. (Caudalejeunea samoana Steph.) from Samoa. Both C. grolleana and C. cristiloba have a 4-5-carinate perianth, which shows that the trigonous perianth present in most species of Caudalejeunea is not a stable character of this genus.

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Journal Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht
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Citation
Gradstein, S.R. (1974). Studies on Lejeuneaceae subf. Ptychanthoideae (Hepaticae) II. Two remarkable species of Caudalejeunea: C. grolleana spec. nov. and C. cristiloba (Steph.) comb. nov. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht, 406(1), 333–343.