Previous molecular phylogenetic studies indicated expansion of Breynia with inclusion of Sauropus s.str. (excluding Synostemon). The present study adds qualitative and quantitative morphological characters to molecular data to find more resolution and/or higher support for the subgroups within Breynia However, the results show that combined molecular and morphological characters provide limited synergy. Morphology confirms and makes the infrageneric groups recognisable within Breynia The status of the Sauropus androgynus complex is discussed. Nomenclatural changes of Sauropus species to Breynia are formalised. The genus Synostemon is reinstated.

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Blumea: Biodiversity, Evolution and Biogeography of Plants

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van Welzen, P. C., Pruesapan, K., Telford, I. R. H., Esser, H.-J., & Bruhl, J. J. (2014). Phylogenetic reconstruction prompts taxonomic changes in Sauropus, Synostemon and Breynia (Phyllanthaceae tribe Phyllantheae). Blumea: Biodiversity, Evolution and Biogeography of Plants, 59(2), 77–94.