A contribution to the intraspecific systematics of Bufo bufo (Linnaeus, 1758) (Amphibia)
Beaufortia , Volume 21 - Issue 280 p. 99- 116
A biometrical analysis of preserved specimens of Bufo bufo, supplemented by a detailed examination of the shape of the skin warts, was performed in order to ascertain the taxonomic status of the form described as B.b.spinosus. Indications of a N.-S. clinal variation in the formation of wart thorns and in allometric relations of body elements were found. The validity of the form as a distinct subspecies seems accordingly very doubtful.
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de Lange, L. (1973). A contribution to the intraspecific systematics of Bufo bufo (Linnaeus, 1758) (Amphibia). Beaufortia, 21(280), 99–116.